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I've also heard these are a good source of tannins, and were used to make the original iron based inks.

Been having fun finding galls - swellings or growths on leaves & stems caused by insects, mites, & even fungi. There are so many different shapes and they're all so fascinating! You can find many on oak and hackberry trees. Just flip over a leaf! #galls…
Small, circular, yellowish to red colored jumping oak galls scattered on the underside of an oak leaf.
Raised bumps known as hackberry nipple gall scattered on the underside of a leaf
Light-green colored cone galls aligned in the center of a witch hazel (Hamamelis) leaf
Pointed-shaped galls known as hackberry thorn gall on the underside of a leaf